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Sannine Formation

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Coordinates: 34°06′N 35°42′E / 34.1°N 35.7°E / 34.1; 35.7
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Sannine Formation
Stratigraphic range: Late Cenomanian
~95–93.5 Ma
↓
Fossil block from Haqel with fishes and a lobster
TypeGeological formation
UnderliesMaameltain Formation
OverliesHammana Formation
ThicknessCoastal: >2,000 m (6,600 ft)
Mountains: 500–700 m (1,600–2,300 ft)
Lithology
PrimaryChalk, limestone
OtherMarl
Location
Coordinates34°06′N 35°42′E / 34.1°N 35.7°E / 34.1; 35.7
Approximate paleocoordinates12°54′N 30°24′E / 12.9°N 30.4°E / 12.9; 30.4
RegionJabal Lubnan
Country Lebanon
Type section
Named forMount Sannine
Sannine Formation is located in Lebanon
Sannine Formation
Sannine Formation (Lebanon)

The Sannine Formation, also called the Sannine Limestone, is a Cretaceous geologic formation in Lebanon. It is a Konservat-Lagerstätte that contains a high diversity of well-preserved fish, reptiles, and invertebrates from the Tethys Ocean within its three main localities: Haqel (alternatively Hakel or Haqil), Hjoula (alternatively Hadjoula, Hajoula, or Hgula), and Nammoura (alternatively Namoura).[1]

It is one of three major Cretaceous lagerstätte in Lebanon, alongside the older (Barremian-aged) Lebanese amber and the younger (Santonian-aged) Sahel Aalma site. The Sannine Formation localities, combined with Sahel Alma, are together referred to as the "Fish Beds" of Lebanon.[1]

Description

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It is primarily Cenomanian in age, with Haqel and Hjoula being late Cenomanian, while the slightly older Nammoura site is middle Cenomanian.[1] Although Lebanon is now a part of Asia, the depositional environment for both formations would have been located off the coast of northern Africa during the Cretaceous, which would technically make their biota African rather than Asian.[2][3]

The Cretaceous formations of Lebanon laterally vary from east to west; the western lowland "coastal" sequence (containing Sahel Aalma) is over 2000 metres thick and primarily consists of deep water limestone and chalk, while the eastern "mountain" sequence (containing the Sannine Formation) is 500–700 metres thick and consists of shallow water limestone.[4] Two different shallow-water environments are preserved by the formation: Nammoura preserves a nearshore inner continental shelf environment with significant terrestrial input, while Haqel & Hjoula preserve an outer continental shelf environment. The neritic nature of these habitats contrasts with the other major marine lagerstätte of the region, Sahel Aalma, which is thought to have been deposited in a deepwater continental slope environment.[1]

Modern position of the Hjoula locality compared to its Cretaceous location.

Pterosaur fossils have been recovered from the formation, including those of Mimodactylus[3] and Microtuban.[5] The pythonomorphs Pontosaurus, Eupodophis and probably Aphanizocnemus are known from the formation.[6] Insects are also known from compression fossils, including those of a dragonfly,[7] enigmatic pond-skater like insect Chresmoda.[8] and a mesoblattinid cockroach.[9] Compression fossils of angiosperm Sapindopsis are also known.[10] Numerous species of fossil fish are known,[11] including the ionoscopid Spathiurus dorsalis[12] and pycnodontid Flagellipinna rhomboides.[13] An incredible diversity of fossil pycnodonts of various, highly unusual body plans is known, in addition to the earliest representatives of modern fish groups such as acanthomorphs, African butterflyfish and eels.[1]

History

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The Sannine Formation and its fossil fishes have been known of since Roman Phoenicia. The earliest mention of them is in the Chronicon by Eusebius around 314 CE (references to earlier mentions by Herodotus are thought to be apocryphal), who mentions fossil fishes commonly being found during quarrying operations in the mountains, and cites them as evidence of the Biblical Flood. The site that these fish were recovered from is thought to correspond to the modern Haqel site. Around 1250, fossil fishes were presented to Louis IX during the Seventh Crusade and documented by Jean de Joinville, although it remains uncertain whether these originated from Haqel or the younger Sahel Alma site.[1][14]

The first European to visit a Lebanese fossil fish locality in person was Cornelis de Bruijn in 1698, who illustrated a fossil fish that corresponds with the species Prionolepis cataphractus from Haqel. The first scientific presentation on the fossil fishes of Haqel was in 1703 at the French Academy of Sciences, and the first scientific studies were conducted by Henri Marie Ducrotay de Blainville in 1818. Lady Hester Stanhope collected and circulated fossil fishes from Haqel and Sahel Alma during the early 19th century, contributing to the study of these specimens by scientists such as Louis Agassiz. The Hadjoula site was only discovered in the late 19th century, and the Nammoura site in the early 20th century.[1][14]

Paleobiota

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Vertebrates

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Jawless fish

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Agnathans reported from the Sannine Formation
Genus Species Presence Notes Images
Tethymyxine T. tapirostrum Hjoula One of the earliest known definitive fossil hagfish.[15]

Cartilaginous fish

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Chondrichthyans reported from the Sannine Formation[16][17]
Genus Species Presence Notes Images
Cantioscyllium C. decipiens Haqel & Hjoula A nurse shark.
C. sp. Hjoula
Crassescyliorhinus C. cf. germanicus Hjoula A catshark.
Cretalamna C. sp. Hjoula An otodontid shark, formerly identified as C. appendiculata.[18]
Cretodus C. "longiplicatus" Hjoula A pseudoscapanorhynchid shark.
C. semiplicatus Haqel & Hjoula
Cretoxyrhina C. mantelli Hjoula A ginsu shark.
Cyclobatis C. major Haqel & Hjoula A cyclobatid ray.[19][20]
C. oligodactylus Haqel, Hjoula & Nammoura[20]
C. tuberculatus Haqel
Deania D. sp. Haqel A gulper shark.
Ginglymostomatidae indet. Hjoula A nurse shark.
Haimirichia H. amonensis Haqel & Nammoura A haimirichiid mackerel shark.
Hemiscylliidae indet. Haqel & Hjoula A bamboo shark.
Heterodontus H. sp. Hjoula A bullhead shark.
Libanopristis L. hiram Haqel & Hjoula A sawskate.
Mesiteia M. emiliae Haqel A mesiteiid carpet shark.
Micropristis M. solomonis Hjoula A sawskate.
Odontaspididae indet. Hjoula A sand shark.
Odontaspis O. aculeatus Hjoula A sand shark.
O. sp.
Parascyllidae indet. Haqel & Hjoula A collared carpet shark.
Paranomotodon P. sp. Hjoula A thresher shark.
Protoscyliorhinus P. sp. Haqel A catshark.
Pseudocorax P. kindlimmani Haqel A false crow shark.[21]
Rajorhina R. expansa Hjoula A skate.
'Rhinobatos' (gen. nov. 1) R. grandis Haqel A potential relative of wedgefishes. Likely its own genus.[22]
R. maronita
'Rhinobatos' (gen. nov. 3) R. hakelensis Haqel A guitarfish-like ray. Likely its own genus, and its status as a guitarfish is disputed.[22]
R. whitfieldi Hjoula
Rhombopterygia R. rajoides Hjoula A guitarfish.
Scyliorhinidae indet. Haqel A catshark.
Scyliorhinus S. arambourgi Hjoula A catshark.
S. bloti Haqel
S. sp.
Squalicorax S. falcatus Nammoura A crow shark.
S. sp. Haqel, Hjoula & Nammoura
Squatina S. sp. Haqel An angelshark.
Triakidae indet. Haqel A houndshark.

Ray-finned fish

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Actinopterygians reported from the Sannine Formation[16]
Genus Species Presence Notes Images
Abisaadia A. hakelensis Haqel An eel.
Abisaadichthys A. libanicus Hjoula A protobramid tselfatiiform.
Acrognathus A. dodgei Haqel & Hjoula A greeneye.
Acrorhinichthys A. poyatoi Haqel & Nammoura A pycnodont of uncertain affinities.
Aipichthys A. minor Haqel & Hjoula A lamprimorph.
A. oblongus Haqel
A. velifer Haqel
Anguillavus A. mazeni Haqel & Hjoula An eel.
A. quadripinnis Hjoula
Apateopholidae indet. nov. gen. nov. sp. Haqel An apatepopholid aulopiform.
Apatepopholis A. laniatus Haqel, Hjoula & Nammoura An apatepopholid aulopiform.
A. nov. sp. Nammoura
Aphanepygus A. dorsalis Haqel An aphanepygid.
Akromystax A. tilmachiton Haqel & Nammoura A pycnodont of uncertain affinities.
Armigatus A. alticorpus Nammoura[23] and Haqel[24] A clupeomorph[24][23]
A. brevissimus Haqel and Hjoula[24] An armigatid clupeomorph[24]
A. namourensis Nammoura[23] A clupeomorph[24]
Belonostomus B. "sp. 1" Nammoura[23] An aspidorhynchid. 2 unnamed species awaiting better material[23]
B. "sp. 2"
Benthesikyme B. serpentina Haqel & Hjoula A dercetid aulopiform.[25]
Berycopsis B. pulcher Hjoula A beardfish.
Capassopiscis C. pankowskii Haqel A pantodontid.[26]
Charitopsis C. spinosus Haqel & Hjoula A gonorynchid.
Chirocentrites C. sp. Nammoura A cladocyclid ichthyodectiform.
Coccodus C. armatus Haqel A coccodontid pycnodont.
C. insignis Hjoula
Corusichthys C. megacephalus Haqel A coccodontid pycnodont.
Cretapantodon C. polli Haqel A pantodontid.[27]
Cryptoberyx C. minimus Hjoula An indeterminate acanthopterygian.
Ctenodentelops C. striatus Nammoura[23] An elopid.[23]
Ctenothrissa C. protodorsalis Haqel A ctenothrissiform acanthomorph.
C. signifer Hjoula
C. vexillifer Haqel
?Cylindracanthus ?C. libanicus Haqel & Hjoula An indeterminate ray-finned fish known only from rostral spines.
Davichthys D. gardnieri Haqel An elopid.
Dercetidae indet. nov. gen. nov. sp. Hjoula A dercetid aulopiform.
Diplomystus D. birdi Haqel & Hjoula An armigatid clupeomorph.
Ducrotayichthys D. cornutus Haqel A gladiopycnodontid pycnodont.
Enchelion E. montium Haqel & Hjoula An eel.
Enchodus E. marchesetti Haqel & Hjoula An enchodontid aulopiform.
E. mecoanalis Nammoura[23]
E. "sp. 1." Haqel
E. "sp. 2." Nammoura
Eubiodectes E. libanicus Haqel, Hjoula & Nammoura A cladocyclid ichthyodectiform.
Eurypholidae indet. nov. gen. nov. sp. Nammoura An enchodontid aulopiform.
Eurypholis E. boissieri Haqel, Hjoula & Nammoura An enchodontid aulopiform.
Eusebichthys E. byblosi Haqel A protobramid tselfatiiform.
E. "sp. 1" Haqel
E. "sp. 2" Hjoula
Exocoetoides E. minor Haqel & Hjoula A cheirothricid.
Flagellipinna F. rhomboides Haqel A pycnodontid pycnodont.[13]
Gebrayelichthys G. uyenoi Haqel A gebrayelichthyid pycnodont.
G. vexillarius Haqel
Gaudryella G. gaudryi Haqel, Hjoula & Nammoura An indeterminate euteleost.
Gharbouria G. libanica Nammoura An indeterminate euteleost.
Gigapteryx G. tethyestris Nammoura[23] A lamprimorph.[23][28]
Gladiopycnodus G. byrnei Hjoula A gladiopycnodontid pycnodont.[29]
G. karami Haqel
Ginsburgia G. operta Haqel & Hjoula An indeterminate euteleost.
Hajulia H. multidens Haqel & Hjoula A bonefish.
Hakelia H. laticauda Haqel & Hjoula An indeterminate neoteleost.
Hakeliosomus H. hakelensis Haqel & Hjoula A gonorynchid.
Haqelpycnodus H. picteti Haqel A pycnodontid pycnodont.
Hastichthys H. gracilis Haqel & Nammoura A dercetid aulopiform.
Hayenchelys H. germanus Hjoula An eel.
Hayolperichthys H. pectospinus Haqel A gladiopycnodontid pycnodont.
Hemisaurida H. hakelensis Haqel A halecid aulopiform.
Hensodon H. spinosus Haqel A coccodontid pycnodont[30]
Heterothrissa H. signeuxae Haqel A ctenothrissiform acanthomorph.
Hgulichthys H. spinus Hjoula A trachichthyiform.
Humilichthys H. orientalis Hjoula A ctenothrissiform acanthomorph.
Ichthyoceros I. spinosus Haqel A coccodontid pyncodont.
Ichthyotringa I. delicata Haqel & Hjoula An ichthyotringid aulopiform.
Joinvillichthys J. kriweti Haqel & Hjoula A gladiopycnodontid pycnodont.
J. lindstroemi Haqel
Lebonichthys L. lewisi Hjoula A bonefish.[23]
L. namourensis Nammoura[23]
Lebrunichthys L. nammourensis Nammoura A pachyrhizodontid.
Libanoberyx L. spinosus A trachichthyiform.
Lissoberyx L. arambourgi Hjoula A trachichthyiform.
L. dayi
L. denticulatus
Luenchelys L. minimus Nammoura An eel.
Maraldichthys M. verticalis Haqel A gebrayelichthyid pycnodont.
Microcapros M. libanicus Hjoula A quaesitoberycid beryciform.[31]
Monocerichthys M. scheuchzeri Haqel & Hjoula A gladiopycnodontid pycnodont.
Nematonotus N. bottae Haqel & Nammoura A flagfin.
N. longispinus Hjoula
'Nursallia' 'N.' goedeli Haqel, Hjoula & Nammoura A pycnodontid pycnodont.
'N.' tethysensis Nammoura and Hjoula
Omosoma O. sp./gen. nov. sp. nov. Nammoura Possibly an indeterminate beardfish,[31] potentially a new genus.
Ornategulum O. sardinioides Haqel & Nammoura A clupeomorph.
Palaeopantodon P. vandersypeni Haqel[32] A pantodontid.
Pankowskichthys P. libanicus Haqel & Nammoura A gladiopycnodontid pycnodont.
Pankowskipiscis P. haqelensis Haqel[33] A pantodontid.
Paracentrus P. lebanonensis Nammoura[23] A holocentriform[23]
P. sp. nov.
Paraclupeidae indet. nov. gen. 1 Nammoura A paraclupeid clupeomorph; two different genera.
nov. gen. 2. Hjoula
Paracoccodus P. woodwardi Haqel A coccodontid pyncodont.
Pateroperca P. libanica Hjoula A ctenothrissiform acanthomorph.
P. robusta
Pattersonichthys P. delicatus Hjoula A ctenothrissiform acanthomorph.
Pattersonoberyx P. pharsus Haqel A potential holocentriform.[31]
Petalopteryx P. syriacus Haqel An aphanepygid.
Petersichthys P. libanicus Haqel A pantodontid.
Pharmacichthys P. numismalis Haqel A pharmacichthyid lamprimorph.
P. venenifer
Phoenicolepis P. arcuatus Hjoula A ctenothrissiform acanthomorph.
Phylactocephalus P. microlepis Haqel & Hjoula A halecid aulopiform.
Plectocretacicus P. clarae Haqel A potential tetraodontiform.
Plesioberyx P. discoides Haqel An indeterminate acanthopterygian.
P. maximus
Prognathoglossum P. kalassyi Haqel A pantodontid.
Prionolepis P. cataphractus Haqel & Hjoula A prionolepid aulopiform.
Protobrama P. avus Haqel & Hjoula A protobramid tselfatiiform.
P. woodwardi Haqel
Protobramidae indet. nov. gen. nov. sp. Nammoura A protobramid tselfatiiform.
Pseudoberyx P. bottae Haqel & Hjoula An indeterminate teleost.
P. grandis Haqel
P. syriacus Haqel & Hjoula
Pycnosteroides P. levispinus Hjoula A pycnosteroidid lamprimorph.
Quaesitoberyx Q. minutus Hjoula A quaesitoberycid beryciform.[31]
Rhamphoichthys R. taxidiotis Haqel[34] A billfish-like plethodid tselfatiiform.
Rhinopycnodus R. gabriellae Haqel A pycnodontid pycnodont.
Rhynchodercetis R. hakelensis Haqel & Hjoula A dercetid aulopiform.
Rostropycnodus R. gayeti Haqel A gladiopycnodontid pycnodont.
Sardinioides S. attenuatus Haqel & Hjoula A sardinioidid myctophiform.
S. minimus
S. pontivagus
Saurorhamphus S. giorgiae Nammoura An enchodontid aulopiform.
Scombroclupea S. diminuta Nammoura[23] A clupeomorph.[23]
S. macrophthalma Haqel & Hjoula
Scutatoclupea S. bacchiai Nammoura A paraclupeid clupeomorph.
Sedenhorstia S. dayi Hjoula A tarpon.
S. libanica Haqel
S. orientalis Hjoula
Serrilepis S. minor Nammoura[23] A halecid aulopiform.[23]
S. prymnostrigos
Sigmapycnodus S. giganteus Haqel A pycnodontid pycnodont.
Sorbinichthys S. elusivo Nammoura A sorbinichthyid clupeomorph.
Spathiurus S. dorsalis Haqel An ionoscopiform.[12]
Spinascutichthys S. pankowskiae Nammoura An enchodontoid aulopiform.[35]
Stanhopeichthys S. libanicus Hjoula A pachyrhizodontid.
Stenoprotome S. hamata Haqel A gladiopycnodontid pycnodont.
Stichocentrus S. elegans Hjoula A holocentriform.
S. liratus
S. spinulosus
S. sp.
Stichopteryx[31] S. lewisi Haqel & Hjoula A trachichthyiform.
Telepholis T. tenuis Haqel & Hjoula A cheirothricid.
Trewavasia T. carinata Haqel A coccodontid pycnodont.[30]
Tricerichthys T. wenzi Haqel A gladiopycnodontid pycnodont.
Triplomystus T. noorae Nammoura[23] A paraclupeid clupeomorph.[23]
T. oligoscutatus
Volcichthys V. sp. Haqel An indeterminate aulopiform or sardinioid myctophiform.
Ypsiloichthys Y. sibelleae Haqel[36] A teleost of uncertain affinities.

Lobe-finned fish

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Sarcopterygians reported from the Sannine Formation
Genus Species Presence Notes Images
Macropomoides M. orientalis Haqel & Hjoula A latimeriid coelacanth.

Amniotes

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Amniotes reported from the Sannine Formation
Genus Species Presence Notes Images
Aphanizocnemus A. libanensis Although the type locality is unknown, it is said to "almost certainly" originate from the Sannine Formation. A marine lizard.[6]
Enantiophoenix E. electrophyla Nammoura An enantiornithine bird.[37]
Eupodophis E. descouensi Nammoura An early marine simoliophiid snake with hind limbs.[6]
Microtuban M. altivolans Hjoula An azdarchoid pterosaur.[5]
Mimodactylus M. libanensis Hjoula An istiodactyliform pterosaur.[3]
Pontosaurus P. kornhuberi Nammoura A marine dolichosaurid lizard.[6]
Rhinochelys R. nammourensis Nammoura A marine protostegid turtle. Adult and multiple hatchling remains known, suggesting a nesting site was nearby.[38]
Theropoda indet. Nammoura The partial arm of a theropod dinosaur, the only non-avialan dinosaur material known from the formation. Now lost.[1]

Invertebrates

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Arthropods

[edit]
Arthropods reported from the Sannine Formation[39]
Genus Species Presence Notes Images
Acanthochirana A. smithwoodwardi Haqel & Hjoula An aegerid prawn.
Acanthophoenicides A. peterpani A slipper lobster.
Aeger A. libanensis Haqel & Hjoula An aegerid prawn.
Armacrangon A. edwardsi A glyphocrangonid shrimp.
?Bentheogennema B. phoenicia A benthesicymid crustacean.
Callianassa C. sp. Hjoula A callianassid crustacean.[40]
Calliaxina C. hadjoulaensis Hjoula A callianassid crustacean.
Carinacaris C. antennae A penaeid prawn.
C. teruzzi Haqel & Hjoula
Carpopenaeus C. callirostris Haqel & Hjoula A carpopenaeid prawn.
C. septemspinatus Haqel & Hjoula
Charbelicaris C. maronites Hjoula A slipper lobster.[41]
Chresmoda C. libanica Nammoura A water strider-like chresmodid insect.[8]
Corazzatocarcinus C. hadjoulae Hjoula A necrocarcinid crab.[42]
Cretaxiopsis C. libanotica An axiid crustacean.
Cretacalliax C. levantina Hjoula A callianopsid ghost shrimp.[43]
Epipenaeus E. abisaadorum An carpopenaeid shrimp.
Eryma E. oscari Hjoula An erymid "lobster".
Gladicrangon G. coriacea A glyphocrangonid shrimp.
Globulacaris G. garassinoi Haqel A protozoeid thylacocephalan.
Glyphea G. damesi Hjoula A glypheoid crustacean.[40]
Hakelocaris H. vavassorii Haqel & Hjoula A penaeid prawn.
Homolopsis H. aff. edwardsii Hjoula A carrier crab.[42]
Jasus J. sp. Hjoula A spiny lobster.[40]
Joinvilleicaris J. longirostris A penaeid prawn.
Libanoaxius L. beatae An axiid crustacean.
Libanocaris L. curvirostra A penaeid prawn.
L. rogeri Haqel & Hjoula
Lebanoaktassia L. curiosa Hjoula An aktassiid dragonfly.[44]
Libanocordulia L. debiei Hjoula A libanocorduliid dragonfly[7]
Libanogomphus L. lionelcavini Hjoula A libanogomphid dragonfly.[45]
Libanoliupanshania L. mimi Hjoula A liupanshaniid dragonfly[7]
Linuparus L. secretanae A spiny lobster.
Macropenaeus M. incertus Haqel & Hjoula A penaeid prawn.
Magila M. cretacica An axiid crustacean.
Microchela M. rostrata Haqel & Hjoula A penaeid prawn.
Mieroblattina M. pacis Nammoura A mesoblattinid cockroach.[9]
Notahomarus N. hakelensis Haqel & Hjoula A true lobster.
Odontochelion O. bicarinatum A crangonid shrimp.
O. cretaceum Haqel & Hjoula
Palaeobenthonectes P. arambourgi A benthesicymid crustacean.
Palaeopalinurellus P. jbeilensis Haqel A slipper lobster.[46]
Palaeopandalus P. komaii A pandalid shrimp.
Palibacus P. praecursor Haqel & Hjoula A slipper lobster.
Palinurus P. teruzzii A spiny lobster.
Paracancrinos P. libanensis Hjoula A slipper lobster.[41]
Paradollocaris P. vannieri Hjoula A dollocaridid thylacocephalan.
Paranecrocarcinus P. cf. milbournei Hjoula A necrocarcinid crab. Specimen consumed by a catshark.[47]
Phalangiopsis P. antoinei A chirostylid squat lobster.
P. rogeri
Phoenice P. pasinii Haqel A boxer shrimp.[40]
Pseudastacus P. dubertreti Haqel & Hjoula A stenochirid "lobster".
Pustulina P. cretacea[48] Haqel & Hjoula An erymid "lobster".
Striadiogenes S. frigerioi Hjoula A diogenid hermit crab.[49]
Tachypleus T. syriacus Hjoula A horseshoe crab.[50]
Telamonocarcinus T. gambulatus Haqel & Hjoula A dorippid crab.[42]
Thylacocaris T. schrami Hjoula A dollocaridid thylacocephalan.

Molluscs

[edit]
Molluscs reported from the Sannine Formation
Genus Species Presence Notes Images
Allocrioceras A. cf. annulatum An anisoceratid ammonite.[21][51]
Boreopeltis B. smithi Haqel or Hjoula A plesioteuthid coleoid.[52]
Dorateuthis D. syriaca Haqel & Hjoula A plesioteuthid coleoid.[53]
Edmunditeuthis E. bacchiai Haqel & Hjoula A trachyteuthid coleoid.[54][55]
Glyphidopsis G. waagei Haqel A trachyteuthid coleoid.[52]
Glyphiteuthis G. abisaadorum Haqel & Hjoula A trachyteuthid coleoid.[52][56]
G. freijii Haqel, Hjoula & Nammoura
G. libanotica Haqel & Hjoula
Keuppia K. levante Hjoula An early octopus. 2 or 3 species known from the same locality[57]
K. hyperbolaris
K. sp.
Mantelliceras M. mantelli Haqel "the occurrence of Mantelliceras mantelli is regarded doubtful, as this species has been used for various taxa within the family Acanthoceratidae de Grossouvre, 1894, and specimens have never been figured"[21][51]
Rachiteuthis R. acutali Hjoula A palaeololiginid coleoid.[58]
R. donovani Hâqel
Styletoctopus S. annae Hâqel An early octopus.[57]
Teudopsinia T. haasi Haqel A teudopsid coleoid.[54]
Syrionautilus S. libanoticus Hjoula A cymatoceratid nautiloid.[59]

Echinoderms

[edit]
Echinoderms reported from the Sannine Formation[60]
Genus Species Presence Notes Images
Antedon A. pinnulata Haqel A crinoid.
Aspidochirotida indet. Hjoula A sea cucumber.[61]
Geocoma G. libanotica Haqel A brittle star.

Annelids

[edit]
Annelids reported from the Sannine Formation[62]
Genus Species Presence Notes Images
Didone D. pulcherrima A didonid bristleworm.
Eunicites E. diopatroides A eunicid bristleworm.
E. falcatus
E. joinvillei
E. mariacristinae
Ferragutia F. cenomaniana A bristleworm.
Lumbriconereites L. garassinoi A bristleworm.
L. hadjulae
Palaeoaphrodite P. libanica An aphroditid bristleworm.
Phoeniciarabella P. caesaris A lumbrinerid bristleworm.
P. orensanzi
P. pinnulata
Teruzzia T. gryphoeides A lumbrinerid bristleworm.
T. pezzolii
T. pusilla
T. sagittifera
Rollinschaeta R. myoplena Haqel & Hjoula A fireworm.[63]

Plants

[edit]
Plants reported from the Sannine Formation
Genus Species Presence Notes Images
Aryskumia cf. A. zelkovifolia Nammoura Dicot known from a poorly preserved leaf with ovate blade.[64]
Nammouria N. gracilis An aquatic fern of uncertain affinities.[64]
Nammourophyllum N. altingioides A dicot known from leaves, a possible member of the Altingiaceae.[64]
Nupharanthus N. cretacea A dicot known by a preserved flower, possibly a member of the Nympheales.[64]
Parvileguminophyllum P. sp. Dicot known from a complete leaflet, possibly a member of the Fabaceae.[64]
'Phragmites' P. sp. A monocot of uncertain affinities, vaguely similar to the modern reed.[64]
Platycaryeae gen. et sp. indet. Infructescence of bracteate fruits resembling Platycarya americana.[64]
Pseudolarix P. sp. A conifer related to the modern golden larch.[64]
Pseudotorellia P. sp. A gymnosperm of uncertain affinities.[64]
Pteris P. sp. A pteridacean fern.[64]
Sapindopsis S. anhouryi A platanacean, related to the modern plane trees.[64]
S. libanensis

See also

[edit]

References

[edit]
  1. ^ a b c d e f g h George, Hady; Bazzi, Mohamad; Hossny, Tamara El; Ashraf, Nida; Saad, Pierre Abi; Clements, Thomas (2024-04-29). "The famous fish beds of Lebanon: the Upper Cretaceous Lagerstätten of Haqel, Hjoula, Nammoura, and Sahel Aalma". Journal of the Geological Society. 181 (5). doi:10.1144/jgs2023-210. ISSN 0016-7649.
  2. ^ López-Arbarello, Adriana. "The record of Mesozoic fishes from Gondwana (excluding India and Madagascar)". Mesozoic Fishes 3 – Systematics, Paleoenvironments and Biodiversity.
  3. ^ a b c Kellner, Alexander W. A.; Caldwell, Michael W.; Holgado, Borja; Vecchia, Fabio M. Dalla; Nohra, Roy; Sayão, Juliana M.; Currie, Philip J. (2019-11-29). "First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity". Scientific Reports. 9 (1): 17875. Bibcode:2019NatSR...917875K. doi:10.1038/s41598-019-54042-z. ISSN 2045-2322. PMC 6884559. PMID 31784545.
  4. ^ Walley, Christopher D. (1997). The Lithostratigraphy of Lebanon: A Review (PDF). pp. 1–20.
  5. ^ a b Elgin, Ross A.; Frey, Eberhard (December 2011). "A new azhdarchoid pterosaur from the Cenomanian (Late Cretaceous) of Lebanon". Swiss Journal of Geosciences. 104 (S1): 21–33. doi:10.1007/s00015-011-0081-1. ISSN 1661-8726. S2CID 128405107.
  6. ^ a b c d Campbell Mekarski, Michelle; Pierce, Stephanie E.; Caldwell, Michael W. (2019-09-24). "Spatiotemporal Distributions of Non-ophidian Ophidiomorphs, With Implications for Their Origin, Radiation, and Extinction". Frontiers in Earth Science. 7: 245. Bibcode:2019FrEaS...7..245M. doi:10.3389/feart.2019.00245. ISSN 2296-6463.
  7. ^ a b c Azar, Dany; Maksoud, Sibelle; Huang, Diying; Nel, André (January 2019). "First Lebanese dragonflies (Insecta: Odonata, Aeshnoptera, Cavilabiata) from the Arabo-African mid-Cretaceous paleocontinent" (PDF). Cretaceous Research. 93: 78–89. Bibcode:2019CrRes..93...78A. doi:10.1016/j.cretres.2018.08.025. S2CID 134308438.
  8. ^ a b Nel, Andre; Azar, Dany; Martinez-Delclos, Xavier; Makhoul, Edouard (2004-03-25). "A new Upper Cretaceous species of Chresmoda from Lebanon - a latest representative of Chresmodidae (Insecta: Polyneoptera inc. sed.): first record of homeotic mutations in the fossil record of insects". European Journal of Entomology. 101 (1): 145–151. doi:10.14411/eje.2004.019.
  9. ^ a b P. Vršanský and E. Makhoul. 2013. Mieroblattina pacis gen. et sp. n. - Upper Cretaceous cockroach (Blattida: Mesoblattinidae) from Nammoura limestone of Lebanon. Insect Evolution in an Ambiferous and Stone Alphabet 167-172
  10. ^ D. L. Dilcher and P. W. Basson. 1990. Mid-Cretaceous angiosperm leaves from a new fossil locality in Lebanon. Botanical Gazette 151(4):538-547
  11. ^ Forey, Peter L.; Yi, Lu; Patterson, Colin; Davies, Cliff E. (January 2003). "Fossil fishes from the Cenomanian (Upper Cretaceous) of Namoura, Lebanon". Journal of Systematic Palaeontology. 1 (4): 227–330. Bibcode:2003JSPal...1..227F. doi:10.1017/S147720190300107X. ISSN 1477-2019. S2CID 85606708.
  12. ^ a b El Hossny, Tamara; Maksoud, Sibelle; Azar, Dany; Saad, Pierre Abi; Cavin, Lionel (August 2020). "Spathiurus dorsalis Davis, 1887, from the Upper Cretaceous of Haqel, Lebanon, and the evolutionary history of Ionoscopiformes". Cretaceous Research. 116: 104619. Bibcode:2020CrRes.11604619E. doi:10.1016/j.cretres.2020.104619. S2CID 224978290.
  13. ^ a b Cawley, John Joseph; Kriwet, Jürgen (2019-03-04). "A new genus and species of pycnodontid fish Flagellipinna rhomboides , gen. et sp. nov. (Neopterygii, Pycnodontiformes), from the Upper Cretaceous (Cenomanian) of Lebanon, with notes on juvenile form and ecology". Journal of Vertebrate Paleontology. 39 (2): e1614012. Bibcode:2019JVPal..39E4012C. doi:10.1080/02724634.2019.1614012. ISSN 0272-4634. PMC 6817322. PMID 31709027.
  14. ^ a b Capasso, Luigi (2017). "The history and the situation of the world famous fossil fish quarries in Lebanon". Bollettino del Museo Civico di Storia Naturale di Verona. 41: 53–76.
  15. ^ Miyashita, Tetsuto; Coates, Michael I.; Farrar, Robert; Larson, Peter; Manning, Phillip L.; Wogelius, Roy A.; Edwards, Nicholas P.; Anné, Jennifer; Bergmann, Uwe; Palmer, A. Richard; Currie, Philip J. (2019-02-05). "Hagfish from the Cretaceous Tethys Sea and a reconciliation of the morphological–molecular conflict in early vertebrate phylogeny". Proceedings of the National Academy of Sciences. 116 (6): 2146–2151. Bibcode:2019PNAS..116.2146M. doi:10.1073/pnas.1814794116. ISSN 0027-8424. PMC 6369785. PMID 30670644.
  16. ^ a b Amalfitano, Jacopo; Giusberti, Luca; Fornaciari, Eliana; Carnevale, Giorgio (2020-04-03). "UPPER CENOMANIAN FISHES FROM THE BONARELLI LEVEL (OAE2) OF NORTHEASTERN ITALY". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2). doi:10.13130/2039-4942/13224. ISSN 2039-4942.
  17. ^ Pradel, Alan; Denton, John S. S.; Janvier, Philippe; Maisey, John G., eds. (2021). Ancient fishes and their living relatives: a tribute to John G. Maisey. München: Verlag Dr. Friedrich Pfeil. ISBN 978-3-89937-269-4.
  18. ^ Greenfield, T. (2022). "Additions to "List of skeletal material from megatooth sharks", with a response to Shimada (2022)". Paleoichthys. 6: 6–11.
  19. ^ Egerton, Philip Grey (1845). "Description of a Fossil Ray from Mount Lebanon (Cyclobatis oligodactylus)". Quarterly Journal of the Geological Society. 1 (1): 225–229. doi:10.1144/GSL.JGS.1845.001.01.60. S2CID 128842555.
  20. ^ a b "A new Miocene skate from the Central Paratethys (Upper Austria): the first unambiguous skeletal record for the Rajiformes (Chondrichthyes: Batomorphii) (project)". MorphoBank datasets. 2019-04-29. Retrieved 2022-07-22.
  21. ^ a b c Jambura; Stumpf; Kriwet, Patrick L.; Sebastian; Jürgen (September 2021). "Skeletal remains of the oldest known pseudocoracid shark Pseudocorax kindlimanni sp. nov. (Chondrichthyes, Lamniformes) from the Late Cretaceous of Lebanon". Cretaceous Research. 125: 104842. Bibcode:2021CrRes.12504842J. doi:10.1016/j.cretres.2021.104842. PMC 7611798. PMID 34642522.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  22. ^ a b Georges Kachacha, Gilles Cuny (2017). "Revision of the fossil batomorphs from the Cretaceous of Lebanon, and their impact on our understanding of the early step of the evolution of the clade". Research & Knowledge. 3: 2. doi:10.14456/RANDK.2017.22.
  23. ^ a b c d e f g h i j k l m n o p q r s t 1945-, Forey, Peter L. (Peter Lawrence), (2003). Fossil fishes from the Cenomanian (Upper Cretaceous) of Namoura, Lebanon. Cambridge University Press. OCLC 66582935.{{cite book}}: CS1 maint: extra punctuation (link) CS1 maint: numeric names: authors list (link)
  24. ^ a b c d e Than-Marchese, Bruno Andrés; Alvarado-Ortega, Jesús (2022-10-01). "Armigatus felixi sp. nov. An Albian double armored herring (Clupeomorpha, Ellimmichthyiformes) from the Tlayúa lagerstätte, Mexico". Journal of South American Earth Sciences. 118: 103905. Bibcode:2022JSAES.11803905T. doi:10.1016/j.jsames.2022.103905. ISSN 0895-9811.
  25. ^ Alvarado-Ortega, Jesús; Díaz-Cruz, Jesús Alberto (2021-01-01). "Hastichthys totonacus sp. nov., a North American Turonian dercetid fish (Teleostei, Aulopiformes) from the Huehuetla quarry, Puebla, Mexico". Journal of South American Earth Sciences. 105: 102900. Bibcode:2021JSAES.10502900A. doi:10.1016/j.jsames.2020.102900. ISSN 0895-9811.
  26. ^ Taverne, L. (2022). "The Pantodontidae (Teleostei, Osteoglossomorpha) from the marine Cenomanian (Upper Cretaceous) of Lebanon. 4°. Capassopiscis pankowskii gen. and sp. nov" (PDF). Geo-Eco-Trop. 46 (1): 149–158.
  27. ^ Taverne, L. (2022). "The Pantodontidae (Teleostei, Osteoglossomorpha) from the marine Cenomanian (Upper Cretaceous) of Lebanon. 6°. Cretapantodon polli gen. and sp. nov" (PDF). Geo-Eco-Trop. 46 (1): 149–158.
  28. ^ Delbarre, Daniel J.; Davesne, Donald; Friedman, Matt (2016-07-02). "Anatomy and relationships of † Aipichthys pretiosus and †' Aipichthys ' nuchalis (Acanthomorpha: Lampridomorpha), with a review of Late Cretaceous relatives of oarfishes and their allies". Journal of Systematic Palaeontology. 14 (7): 545–567. Bibcode:2016JSPal..14..545D. doi:10.1080/14772019.2015.1078538. ISSN 1477-2019.
  29. ^ Marramà, Giuseppe; Villier, Boris; Dalla Vecchia, Fabio M.; Carnevale, Giorgio (2016-06). "A new species of Gladiopycnodus (Coccodontoidea, Pycnodontomorpha) from the Cretaceous of Lebanon provides new insights about the morphological diversification of pycnodont fishes through time". Cretaceous Research. 61: 34–43. doi:10.1016/j.cretres.2015.12.022. ISSN 0195-6671.
  30. ^ a b Kriwet, Jürgen (2004-09-10). "A new pycnodont fish genus (Neopterygii: Pycnodontiformes) from the Cenomanian (Upper Cretaceous) of Mount Lebanon". Journal of Vertebrate Paleontology. 24 (3): 525–532. doi:10.1671/0272-4634(2004)024[0525:anpfgn]2.0.co;2. ISSN 0272-4634. S2CID 129294502.
  31. ^ a b c d e Murray, Alison M. (2014). "Mid-Cretaceous acanthomorph fishes with the description of a new species from the Turonian of Lac des Bois, Northwest Territories, Canada". Vertebrate Anatomy Morphology Palaeontology. 1: 101–115. doi:10.18435/B5CC78. ISSN 2292-1389.
  32. ^ Taverne, L. (2021). "The Pantodontidae (Teleostei, Osteoglossomorpha) from the marine Cenomanian (Upper Cretaceous) of Lebanon. 3°. Palaeopantodon vandersypeni gen. and sp. nov" (PDF). Geo-Eco-Trop. 45 (4): 699–708.
  33. ^ Taverne, L. (2021). "The Pantodontidae (Teleostei, Osteoglossomorpha) from the marine Cenomanian (Upper Cretaceous) of Lebanon. 1°. Pankowskipiscis haqelensis gen. and sp. nov" (PDF). Geo-Eco-Trop. 45 (4): 551–562.
  34. ^ El Hossny, Tamara; Cavin, Lionel; Kaplan, Ulrich; Schwermann, Achim H.; Samankassou, Elias; Friedman, Matt (2023). "The first articulated skeletons of enigmatic Late Cretaceous billfish-like actinopterygians". Royal Society Open Science. 10 (12): 231296. Bibcode:2023RSOS...1031296E. doi:10.1098/rsos.231296. ISSN 2054-5703. PMC 10698480. PMID 38077217.
  35. ^ Murray, Alison M.; Chida, Mori; Holmes, Robert B. (2022-06-30). "New enchodontoid fish (Teleostei: Aulopiformes) from the Late Cretaceous of Lebanon". Journal of Vertebrate Paleontology. 42 (1). Bibcode:2022JVPal..42E1370M. doi:10.1080/02724634.2022.2101370. ISSN 0272-4634.
  36. ^ El Hossny, Tamara; Cavin, Lionel (July 2023). "A New Enigmatic Teleost Fish from the Mid-Cretaceous of Lebanon". Diversity. 15 (7): 839. doi:10.3390/d15070839. ISSN 1424-2818.
  37. ^ Cau, A.; Arduini, P. (2008). "Enantiophoenix electrophyla gen. et sp. nov. (Aves, Enantiornithes) from the Upper Cretaceous (Cenomanian) of Lebanon and its phylogenetic relationships". Atti della Societa Italiana di Scienze Naturali e del Museo Ivico di Storia Naturale in Milano. 149 (2): 293–324.
  38. ^ Rhinochelys (Chelonioidea: Protostegidae) from the Late Cretaceous (Cenomanian) of Nammoura, Lebanon
  39. ^ Sylvain Charbonnier; Denis Audo; Alessandro Garassino; Matúš Hyžný (2017). Fossil Crustacea of Lebanon. National Museum of Natural History. pp. 1–252. ISBN 978-2-85653-785-5.
  40. ^ a b c d Garassino, A. (2000). "New decapod crustaceans from the Cenomanian (Upper Cretaceous) of Lebanon". Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano. S2CID 88896292.
  41. ^ a b Haug, Joachim T.; Audo, Denis; Charbonnier, Sylvain; Palero, Ferran; Petit, Gilles; Abi Saad, Pierre; Haug, Carolin (2016-03-01). "The evolution of a key character, or how to evolve a slipper lobster". Arthropod Structure & Development. Special Issue: Fossils as Living Beings. 45 (2): 97–107. Bibcode:2016ArtSD..45...97H. doi:10.1016/j.asd.2015.08.003. ISSN 1467-8039. PMID 26319267.
  42. ^ a b c Larghi, Cristiano (2004). "Brachyuran decapod Crustacea from the Upper Cretaceous of Lebanon". Journal of Paleontology. 78 (3): 528–541. Bibcode:2004JPal...78..528L. doi:10.1666/0022-3360(2004)078<0528:BDCFTU>2.0.CO;2. ISSN 0022-3360.
  43. ^ Pasini, Giovanni; Poore, Gary C. B.; Garassino, Alessandro (2020-08-28). "A new ghost shrimp (Axiidea, Callianopsidae) from the Late Cretaceous (Cenomanian) of Hadjoula, Lebanon". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 297 (2): 217–225. doi:10.1127/njgpa/2020/0922.
  44. ^ Azar, Dany; Nel, André (2024). "The youngest and first Gondwanan representative of the anisopteran family Aktassiidae from the Upper Cretaceous of Lebanon (Odonata, Petalurida)". Cretaceous Research. 154: 105744. Bibcode:2024CrRes.15405744A. doi:10.1016/j.cretres.2023.105744. ISSN 0195-6671.
  45. ^ Azar, Dany; Nel, André (2023-08-01). "Libanogomphidae, a new extraordinary dragonfly family from the Upper Cretaceous of Lebanon (Odonata, Anisoptera)". Cretaceous Research. 148: 105501. Bibcode:2023CrRes.14805501A. doi:10.1016/j.cretres.2023.105501. ISSN 0195-6671.
  46. ^ Garassino, Alessandro; Pasini, Giovanni (2020-10-02). "A new furry lobster (Crustacea, Decapoda, Synaxidae) from the Upper Cretaceous (Cenomanian) of Lebanon". Palaeodiversity. 13 (1). doi:10.18476/pale.v13.a10. ISSN 1867-6294.
  47. ^ Pasini, Giovanni; Garassino, Alessandro (2011). "Predation on brachyuran (Crustacea, Decapoda): an [sic] unique direct evidence from the Late Cretaceous (Cenomanian) of Hadjoula (Lebanon)". Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano. 152 (II): 233–245.
  48. ^ Devillez, Julien; Charbonnier, Sylvain (2022-03-03). "A synthesis of the evolutionary history of erymoid lobsters (Crustacea, Decapoda, Erymoidea)". Geodiversitas. 44 (9). doi:10.5252/geodiversitas2022v44a9. ISSN 1280-9659.
  49. ^ Garassino, A.; Angeli, Antonio De; Pasini, Giovanni (2009). "A new hermit crab (Crustacea, Anomura, Paguroidea) from the Late Cretaceous (Cenomanian) of Lebanon". Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano. 150 (II): 215–228. S2CID 130639300.
  50. ^ Lamsdell, James C.; McKenzie, Scott C. (2015-12-01). "Tachypleus syriacus (Woodward)—a sexually dimorphic Cretaceous crown limulid reveals underestimated horseshoe crab divergence times". Organisms Diversity & Evolution. 15 (4): 681–693. Bibcode:2015ODivE..15..681L. doi:10.1007/s13127-015-0229-3. ISSN 1618-1077.
  51. ^ a b Wippich, Max G. E.; Lehmann, Jens (September 2004). "Allocrioceras from the Cenomanian (mid-Cretaceous) of the Lebanon and its bearing on the palaeobiological interpretation of heteromorphic ammonites". Palaeontology. 47 (5): 1093–1107. Bibcode:2004Palgy..47.1093W. doi:10.1111/j.0031-0239.2004.00408.x. ISSN 0031-0239. S2CID 129293243.
  52. ^ a b c Fuchs; Larson, Dirk; Neal L (March 2011). "Diversity, Morphology, and Phylogeny of Coleoid Cephalopods from the Upper Cretaceous Plattenkalks of Lebanon–Part I: Prototeuthidina". Journal of Paleontology. 85 (2): 234–249.
  53. ^ Jattiot, Romain; Brayard, Arnaud; Fara, Emmanuel; Charbonnier, Sylvain (2015). "Gladius-bearing coleoids from the Upper Cretaceous Lebanese Lagerstätten: diversity, morphology, and phylogenetic implications". Journal of Paleontology. 89 (1): 148–167. Bibcode:2015JPal...89..148J. doi:10.1017/jpa.2014.13. ISSN 0022-3360.
  54. ^ a b Fuchs, Dirk; Larson, Neal (2011). "Diversity, Morphology, and Phylogeny of Coleoid Cephalopods from the Upper Cretaceous Plattenkalks of Lebanon–Part II: Teudopseina". Journal of Paleontology. 85 (5): 815–834. Bibcode:2011JPal...85..815F. doi:10.1666/10-159.1. ISSN 0022-3360.
  55. ^ Whalen, Christopher D.; Landman, Neil H. (2022-03-08). "Fossil coleoid cephalopod from the Mississippian Bear Gulch Lagerstätte sheds light on early vampyropod evolution". Nature Communications. 13 (1): 1107. Bibcode:2022NatCo..13.1107W. doi:10.1038/s41467-022-28333-5. ISSN 2041-1723. PMC 8904582. PMID 35260548.
  56. ^ Fuchs, D.; Weis, R. (2009-08-01). "A new Cenomanian (Late Cretaceous) coleoid (Cephalopoda) from Hâdjoula, Lebanon". Fossil Record. 12 (2): 175–181. Bibcode:2009FossR..12..175F. doi:10.1002/mmng.200900005. ISSN 2193-0066.
  57. ^ a b Fuchs; Bracchi; Weis, Dirk; Giacomo; Robert (31 December 2008). "NEW OCTOPODS (CEPHALOPODA: COLEOIDEA) FROM THE LATE CRETACEOUS (UPPER CENOMANIAN) OF HÂKEL AND HÂDJOULA, LEBANON". Palaeontology. 52 (1): 65–81. doi:10.1111/j.1475-4983.2008.00828.x. S2CID 129082916.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  58. ^ Jattiot, Romain; Brayard, Arnaud; Fara, Emmanuel; Charbonnier, Sylvain (January 2015). "Gladius-bearing coleoids from the Upper Cretaceous Lebanese Lagerstätten: diversity, morphology, and phylogenetic implications". Journal of Paleontology. 89 (1): 148–167. Bibcode:2015JPal...89..148J. doi:10.1017/jpa.2014.13. ISSN 0022-3360.
  59. ^ Klug, Christian; Pohle, Alexander; Roth, Rosemarie; Hoffmann, René; Wani, Ryoji; Tajika, Amane (2021-07-23). "Preservation of nautilid soft parts inside and outside the conch interpreted as central nervous system, eyes, and renal concrements from the Lebanese Cenomanian". Swiss Journal of Palaeontology. 140 (1): 15. Bibcode:2021SwJP..140...15K. doi:10.1186/s13358-021-00229-9. ISSN 1664-2384. PMC 8549922. PMID 34721283.
  60. ^ "Lebanese Fossils". www.fossilmuseum.net. Retrieved 2024-07-10.
  61. ^ Reich, M. (2004-12-15). Nebelsick, J; Heinzeller, T (eds.). "Echinoderms: Munchen: Proceedings of the 11th International Echinoderm Conference, 6-10 October 2003, Munich, Germany". Echinoderms: Milnchen - Heinzeller & Nebelsick (Eds). Taylor & Francis. doi:10.1201/9780203970881. ISBN 978-0-415-36481-2.
  62. ^ "WoRMS - World Register of Marine Species". www.marinespecies.org. Retrieved 2024-07-10.
  63. ^ Parry, Luke A.; Wilson, Paul; Sykes, Dan; Edgecombe, Gregory D.; Vinther, Jakob (2015-11-17). "A new fireworm (Amphinomidae) from the Cretaceous of Lebanon identified from three-dimensionally preserved myoanatomy". BMC Evolutionary Biology. 15 (1): 256. Bibcode:2015BMCEE..15..256P. doi:10.1186/s12862-015-0541-8. ISSN 1471-2148. PMC 4650273. PMID 26577802.
  64. ^ a b c d e f g h i j k Krassilov, Valentin; Bacchia, Flavio (2000-12). "Cenomanian florule of Nammoura, Lebanon". Cretaceous Research. 21 (6): 785–799. doi:10.1006/cres.2000.0229. ISSN 0195-6671.
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